Play Fact Master on Animated Villains
is the villain in
1995. Ratcliffe is not an out-and-out evil villain. But he is an unpleasant, greedy guy who sees the colonies as nothing more than an opportunity to make a fast buck. Voice Artist: David Ogden Stiers
William G. Kaelin Jr, Sir Peter J. Ratcliffe and Gregg L. Semenza
won for work on
For Their Discoveries of How Cells Sense and Adapt to Oxygen Availability.
2019. "They identified molecular machinery that regulates the activity of genes in response to varying levels of oxygen."
Science & Nature
Nobel Prize in Physiology or Medicine